An up-tick is simply an upward movement in the price of a financial  security such as stock price. This rule affects traders of financial securities such as stock options. In the summer of 2007 an interesting event  occurred in the regulation of securities exchanges, specifically the  Securities and Exchange Commission (SEC) eliminated the requirement for  what is called the up-tick rule. Since then, the combination of the  financial crisis coupled with the elimination of the up-tick rule led to  increased scrutiny of the rule despite the SEC's initial reasons for implementing the 2007 amendments to rule SHO.
The elimination of the up-tick rule  profits short-sellers who are locking into sales prices they think will  be higher than the actual future price. Since the up-tick rule required  there to be an upward movement in price(s) before a short sale  can be initiated, the rule made it more difficult for prices to move  downward as fast as they could have the rule not existed. With the  elimination of the up-tick rule, short sellers of financial securities  have the added advantage of downward price movements in the event of a  low buy to sell price ratio.
Illustration of the uptick rule 
To illustrate the above with an example, if you know the price of  100 cattle is $2000.00/head and you want to profit off the price  movements of those cattle, you may enter into a short-sale contract at  the Chicago Mercantile Exchange (CME). The reason you might do this is  because you think sales of beef are on the decline and that will cause  downward price movement on the price of cattle. If no up-tick rule is in  place and lots of other futures traders also think there will be a  downward movement in the price of cattle, then there is one less  obstacle in the way of declining cattle prices. 
Elimination and reinstatement of the uptick rule
The up-tick rule was eliminated from securities regulation after  the bursting of the housing prices bubble had begun and during the early  phase of the credit-crisis. Financial observers  of the up-tick rule removal postulated the move aggravated and  exaggerated the decline in market prices that helped develop the credit  crisis into something worse causing the ensuing financial crises. In  other words, the timing of the up-tick rule elimination might explain  why some IRA's or 401K's might have experienced shocking losses of  value.
Had the up-tick rule been in place during the  credit-crisis, the massive 1-day declines seen in the stock market might  not have been so volatile and steep. In light of mounting evidence and  pressure to reinstate the up-tick rule, both the SEC and some  legislators within the U.S. House of Representatives reconsidered the  idea of reinstating the up-tick rule in the January 2009 bill 302 of the 111th Congress, after the July House of Representatives Bill 6517  appealing to the same thing failed. Similar but not exactly the same  calls for reinstatement of the up-tick rule were designed by  NYSE-Euronext and other stock exchanges in 2009 as reported by Marketwatch.com  in March of 2009. This in theory would add stability to the economy  during periods of economic decline or slowed growth for the same reasons  the up-tick rule was implemented after the great economic depression of  the 1930's.
Summary
The elimination of the up-tick rule  profits short sellers and increases volatility because it is essentially  removing an obstacle to downward price pressure. If everyone can't sell  at once, it leads to less steep declines and more even securities price  movements because the prices have to go up before each short-sell can  be initiated. In other words, for every step back, a step forward must  be taken first necessitating the step back in price movement to be  larger than the step forward. Moreover, if long positions in securities  decide to sell off, the possibility of amplifying this sell off with  short selling becomes more limited because of the up-tick rule. Hence,  elimination of this rule can lead to both unrestrained dumping of long  positions in addition to leveraged selling through short sales  contracts.
Sources:
1.http://www.foxbusiness.com/story/markets/industries/finance/sec-reinstate-uptick-rule-calm-markets/
2.http://www.investopedia.com/terms/u/uptickrule.asp
3.http://www.investopedia.com/terms/s/shortsale.asp
4.http://seekingalpha.com/article/74825-no-uptick-rule-a-convenient-scapegoat
5.http://www.marketwatch.com
6.http://www.govtrack.us/congress/bill.xpd?bill=h111-302
7. http://tinyurl.com/66yetxk (SEC.gov)
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